Hardouinia sp. Haime in d'Archiac & Haime, 1853
(In Manuscript Preparation)
Plate: HaSp-10
Series: Upper Cretaceous Group: Taylor Formation: Anacacho
Locale: Anacacho Mountains County: Uvalde Stage: Campanian
Length: 44.1 mm Width: 47.8 mm Height: 25.9 mm
Specimen in photo: Collection of Frank & Stephen Crane
SYSTEMATIC PALEONTOLOGY – TAXONOMY
Class: Echinoidea Leske, 1778
Subclass: Euechinoidea Bronn, 1860
Infraclass: Irregularia Latreille, 1825
Superorder: Neognathostomata Smith, 1981
Order: Cassiduloida Claus, 1880
Family: Faujasiidae Lambert, 1905
Genus: HARDOUINIA Haime, 1853
Species: Hardouinia sp.
TYPE SPECIES: Hardouinia potosiensis Lambert, 1911
DIAGNOSIS
A shield-shaped Hardouinia from the Anacacho Formation with a tortoise-like profile, open petals, and a supramarginal periproct situated in a sulcus extending to the margin.
STRATIGRAPHICAL OCCURRENCE & DISTRIBUTION
Group: Taylor
Formations: Anacacho Limestone
Stratigraphic Units: Upper Cretaceous, Campanian
Counties: Uvalde County
DESCRIPTION
TOP: Length, width, height: 44.1 mm L × 47.8 mm W × 25.9 mm H
Top shape: Shield shaped from top; test subconical to high domed (Cooke, 1953 - General Genus).
Top domed: Inflates to a point at the apical system (tortoise shaped)
SIDE: Ambitus: 25% from bottom
Side shape: Sloping sides; posterior end truncated
BOTTOM: Bottom shape: Concave
AMB: Amb shape: Petaloid; petals lanceolate (Cooke, 1953 - General Genus).
Amb arrangement: All 5 ambs near equal in size; bottom 2 slightly larger
Amb primary zones: Five petals do not close on either end. Poriferous zones usually nearly closed in the genus (Cooke, 1953).
PORE PAIRS: Shape: Elongated, oblique; outer pores more elongated than inner (User Description).
Conjugate vs Nonconjugate: Conjugate (Cooke, 1953 - General Genus).
TUBERCLES: Primary tubercles: Small on top, larger on bottom (User Description). Scrobiculate (Cooke, 1953 - General Genus).
APICAL SYSTEM: Location: Posterior of center
Genital pores: 4 gonopores
Madreporite: Large (User Description). Central, separating the posterior oculars (Cooke, 1953 - General Genus).
PERIPROCT (ANUS): Shape & Size: Egg shaped, slightly pointed adorally
Location: Supramarginal; above margin, slightly above midpoint. Sits on truncated posterior end
Sulcus: Sulcus runs from periproct to margin
PERISTOME (MOUTH): Size: Very small
Location: Central
Lip: Bourrelets bulging; largest anteriorly, smallest posteriorly
Phyllodes: Large phyllodes present
SPECIMENS STUDIED & CONDITION Specimen in the collection of Frank & Stephen Crane.
ETYMOLOGY
The genus Hardouinia is named in honor of J. L. Hardouin Michelin (1817–1867), a French paleontologist and zoologist.
REMARKS & COMPARISONS
Remarks: This specimen exhibits characteristics typical of Hardouinia clypeus Cooke, 1955, particularly the "shield shape," the supramarginal periproct in a sulcus, and the Anacacho Formation origin. The genus Hardouinia is characterized by a monobasal apical system and a well-developed floscelle (bourrelets and phyllodes).
Comparisons:
(In Manuscript Preparation)
Plate: HaSp-10
Series: Upper Cretaceous Group: Taylor Formation: Anacacho
Locale: Anacacho Mountains County: Uvalde Stage: Campanian
Length: 44.1 mm Width: 47.8 mm Height: 25.9 mm
Specimen in photo: Collection of Frank & Stephen Crane
SYSTEMATIC PALEONTOLOGY – TAXONOMY
Class: Echinoidea Leske, 1778
Subclass: Euechinoidea Bronn, 1860
Infraclass: Irregularia Latreille, 1825
Superorder: Neognathostomata Smith, 1981
Order: Cassiduloida Claus, 1880
Family: Faujasiidae Lambert, 1905
Genus: HARDOUINIA Haime, 1853
Species: Hardouinia sp.
TYPE SPECIES: Hardouinia potosiensis Lambert, 1911
DIAGNOSIS
A shield-shaped Hardouinia from the Anacacho Formation with a tortoise-like profile, open petals, and a supramarginal periproct situated in a sulcus extending to the margin.
STRATIGRAPHICAL OCCURRENCE & DISTRIBUTION
Group: Taylor
Formations: Anacacho Limestone
Stratigraphic Units: Upper Cretaceous, Campanian
Counties: Uvalde County
DESCRIPTION
TOP: Length, width, height: 44.1 mm L × 47.8 mm W × 25.9 mm H
Top shape: Shield shaped from top; test subconical to high domed (Cooke, 1953 - General Genus).
Top domed: Inflates to a point at the apical system (tortoise shaped)
SIDE: Ambitus: 25% from bottom
Side shape: Sloping sides; posterior end truncated
BOTTOM: Bottom shape: Concave
AMB: Amb shape: Petaloid; petals lanceolate (Cooke, 1953 - General Genus).
Amb arrangement: All 5 ambs near equal in size; bottom 2 slightly larger
Amb primary zones: Five petals do not close on either end. Poriferous zones usually nearly closed in the genus (Cooke, 1953).
PORE PAIRS: Shape: Elongated, oblique; outer pores more elongated than inner (User Description).
Conjugate vs Nonconjugate: Conjugate (Cooke, 1953 - General Genus).
TUBERCLES: Primary tubercles: Small on top, larger on bottom (User Description). Scrobiculate (Cooke, 1953 - General Genus).
APICAL SYSTEM: Location: Posterior of center
Genital pores: 4 gonopores
Madreporite: Large (User Description). Central, separating the posterior oculars (Cooke, 1953 - General Genus).
PERIPROCT (ANUS): Shape & Size: Egg shaped, slightly pointed adorally
Location: Supramarginal; above margin, slightly above midpoint. Sits on truncated posterior end
Sulcus: Sulcus runs from periproct to margin
PERISTOME (MOUTH): Size: Very small
Location: Central
Lip: Bourrelets bulging; largest anteriorly, smallest posteriorly
Phyllodes: Large phyllodes present
SPECIMENS STUDIED & CONDITION Specimen in the collection of Frank & Stephen Crane.
ETYMOLOGY
The genus Hardouinia is named in honor of J. L. Hardouin Michelin (1817–1867), a French paleontologist and zoologist.
REMARKS & COMPARISONS
Remarks: This specimen exhibits characteristics typical of Hardouinia clypeus Cooke, 1955, particularly the "shield shape," the supramarginal periproct in a sulcus, and the Anacacho Formation origin. The genus Hardouinia is characterized by a monobasal apical system and a well-developed floscelle (bourrelets and phyllodes).
Comparisons:
- Hardouinia mortonis: Differs from this specimen by having the periproct located high up between the petals in a deep groove, whereas this specimen has the periproct on the posterior truncation near the margin.
- Hardouinia aequorea: Similar in shape but typically has a smooth, less indented margin and a periproct that is flush or slightly sulcate.
